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Blu, Jewel, and their three children leave the comforts of their domesticated life in Rio de Janeiro to find other blue macaws in the Amazon rainforest. Along the way, Blu faces his most formidable adversary—his father-in-law—while dodging the vengeful cockatoo Nigel.
The story picks up with Blu (Jesse Eisenberg) and Jewel (Anne Hathaway) living a comfortable domestic life in the city with their three children. However, their peaceful existence is disrupted when they discover that they may not be the last of their kind. Reports of other Spix's macaws hiding in the Amazon lead the family on an expedition to find them.